D4.2.13 (HL)—Changes during primary succession
Primary succession gradually changes bare surfaces into more complex communities as pioneers form soil, increase biomass, and support food webs.
- Syllabus
- First assessment 2025
- Objective
- D4.2.13
- Level
- HL
Primary succession gradually changes bare surfaces into more complex communities as pioneers form soil, increase biomass, and support food webs.

Coverage 2014–2025 · Updated 16 Jul 2026
Primary succession begins on a surface without developed soil, so pioneer organisms must help create the conditions needed by later communities.
| From pioneer to later stages | General change |
|---|---|
| Plant size and biomass | Increase |
| Amount of primary production | Increase as producer cover develops |
| Species diversity | Increase as more niches form |
| Food-web complexity | Increase as trophic interactions accumulate |
| Nutrient cycling | Becomes larger and more internally recycled as soil and biomass develop |
On fresh volcanic rock, lichens and mosses weather substrate and add organic matter; deeper soil later supports herbs, shrubs and trees.
If soil and a biological legacy remain after disturbance, recovery is secondary rather than primary succession.
This objective is assessed through structured response, commonly using Describe / Explain / Suggest.
Describe / Explain / Suggest / State / Outline / Predict
Build the answer around this relationship: Pioneer organisms colonize bare surfaces and begin changing the abiotic environment.
Forgetting that primary succession begins on bare substrate with little or no soil.
Representative question
Outline the ecological changes that will occur on the island of cooled lava.
soil develops as lava/rock weathers/breaks down/erodes;
organic material/soil accumulates from (autotrophic) bacteria/lichens;
(gross productivity/biomass increase as) small plants are replaced by larger plants;
development of plant communities support higher trophic levels;
more soil allows for detritivores;
succession increases species diversity / climax community established;
HL D4.2 is the long-term change story. Succession begins after new habitat or disturbance, pioneer species modify conditions, primary succession builds soil and complexity, and some systems repeat cycles or are held before climax by disturbance.
HL succession questions focus on community pathways over time. The answer starts from the initial condition, shows how species modify abiotic conditions and interactions, and identifies whether the pathway is primary, cyclical, climax, or arrested by repeated disturbance.