The nitrogen content of a leaf is mainly due to the proteins contained in the chloroplasts. These proteins are either in the thylakoids or in the stroma, where most enzymes are found. The quantity of nitrogen from the thylakoids is directly proportional to the amount of chlorophyll; a ratio of approximately 50 mmol nitrogen : 1 mmol chlorophyll would represent 100 % of the leaf nitrogen content. Scientists hypothesized that the higher leaf percentage nitrogen content resulting from a decrease in light intensity is due mainly to an increase in chlorophyll in many plant species, three of which are represented in the following graph.