HL adds directionality, helix geometry, DNA packaging, and classic evidence. These ideas stay separate: 5′/3′ explains strand direction; purine-pyrimidine pairing explains width; nucleosomes explain packaging; Hershey-Chase and Chargaff explain evidence for DNA and base pairing.
- Directionality: phosphodiester bonds create 5′ and 3′ ends, and DNA strands are antiparallel.
- Helix stability: purine-pyrimidine pairing keeps width constant and hydrogen bonds stabilize base pairs.
- Packaging: DNA wraps around histone octamers and H1 binds linker DNA.
- Evidence: Hershey-Chase supports DNA as genetic material; Chargaff supports complementary pairing.
- Chargaff found purines equal pyrimidines across DNA samples.