A4.1 Evolution and speciation

Evolution and speciation explain how heritable variation, molecular evidence, selection, isolation, and chromosome change produce population divergence and new species over time.

Syllabus
First assessment 2025
Topic
A4.1
Level
SL

Learning objectives

Evolution changes populations across generations

Evolution is the cumulative change in the heritable characteristics of a population across generations. The population evolves when inherited variants change in frequency; an individual does not evolve during its lifetime.

Change Inherited by offspring? Evolutionary meaning
A DNA variant in a reproductive lineage potentially can enter the population gene pool
A stable heritable change in gene regulation sometimes can contribute while it remains transmissible
Larger muscles from training or a stretched body part no an acquired phenotype, not Darwinian evolution

Mutation and recombination generate heritable variation. Selection, genetic drift and gene flow alter which variants are passed on, so the distribution of characteristics in the next generations can differ from the starting population.

Molecular unity and difference reveal evolutionary history

Nearly all cellular life stores heredity in DNA, uses an almost universal genetic code, translates RNA on ribosomes, transfers energy with ATP and shares core metabolic chemistry. This deep unity is evidence of common ancestry.

Observation in homologous sequences Supported inference
many conserved positions inheritance from a shared ancestor
fewer sequence differences between two taxa usually a more recent common ancestor
differences accumulated after a split evidence of divergence, interpreted with the gene and model used
Aligned homologous molecular sequences show conserved positions and substitutions that support a branching evolutionary relationship.

Sequence similarity is evidence, not a complete history by itself. The sequences must be homologous, sampling must be adequate, and different genes or evolutionary models can yield different estimates.

Selective breeding exposes the power of inherited variation

Selective breeding produces visibly different crop forms from a shared wild ancestor by repeatedly choosing heritable variants.
1

A breeding population already contains heritable variation produced by mutation and recombination. The breeder does not create the desired allele by wanting it.

2

Individuals with a desired phenotype are chosen as parents, so alleles contributing to that phenotype are overrepresented among their offspring.

3

Repeating the choice over generations shifts the population mean and can produce striking breeds or crop forms, as in pigeons, Brassica vegetables and maize.

Artificial and natural selection act on inherited variation, but the selecting agent differs: human preference in breeding, and differential survival or reproduction in the environment.

Homology records divergence; analogy records convergence

Pattern Origin Evolutionary process Example
Homologous structures one ancestral structure, modified for different functions divergent evolution pentadactyl forelimbs for grasping, flight, running or swimming
Analogous structures independent structures shaped by similar selection convergent evolution bat and insect wings as flight surfaces
Homologous vertebrate forelimbs retain one ancestral bone plan despite different functions; bat and insect wings have similar functions but different structural origins.

Similarity of function does not establish close ancestry. Classify evolutionary relationships using homologous features and independent molecular evidence; use analogous features to explain how similar niches can produce similar solutions.

Speciation begins when gene flow no longer unifies a population

A once-connected population experiences reduced gene flow, after which separated gene pools diverge into distinct lineages.
1

One ancestral species begins as populations exchanging alleles through reproduction; gene flow tends to keep their gene pools similar.

2

A geographic, ecological, temporal, behavioural or chromosome barrier reduces allele exchange, so the populations no longer evolve as one fully connected gene pool.

3

Mutation, genetic drift and different selection pressures alter allele frequencies independently on each side of the barrier.

4

If reproductive isolation becomes strong enough that the lineages no longer exchange genes and produce fertile offspring, one pre-existing species has split into two or more species.

Isolation changes which variants succeed in each gene pool

1

Chimpanzees occur mainly north of the Congo River and bonobos south of it. A founder population crossing during a lower-water period, followed by renewed river isolation, is one evidence-based explanation for their divergence from a common ancestor.

2

The river restricts movement and mating, so alleles are not continually mixed between the populations. Isolation creates the conditions for divergence; it does not directly specify which traits will evolve.

3

Different environments and social or ecological interactions can favour different heritable variants, while mutation and genetic drift also change each gene pool independently. Over many generations these processes can establish reproductive isolation.

A present-day barrier and a modern distribution support a historical hypothesis but do not by themselves date the split. Fossil, geological and molecular evidence are needed to test when and how isolation occurred.

Summary: match evolutionary evidence to the claim

Evidence What it supports What it does not show alone
molecular universals deep common ancestry the exact branching history
homologous-sequence differences relative relatedness and divergence a conclusion independent of gene choice or model
selective breeding populations respond rapidly to selection on heritable variation that humans direct natural evolution
homologous structures descent with modification identical function
analogous structures convergence under similar selection recent common ancestry

Evolution can change one connected population without creating a new species. Speciation specifically requires a pre-existing species to split: gene flow falls, gene pools diverge independently, and reproductive isolation prevents renewed allele mixing.

Keep the scale explicit: individuals carry variants and reproduce; populations evolve; lineages split into species.

Evolution as change in heritable characteristics

8 marks

Explain how the process of evolution occurs.

Evidence from sequences

6 marks

Discuss how variations in proteins can indicate phylogeny and be used as an evolutionary clock.

Evidence from selective breeding

3 marks

Outline the evidence for evolution provided by selective breeding.

Evidence from homologous structures

4 marks

Describe the evidence for evolution from homologous structures.

Convergent evolution

4 marks

Explain how analogous structures can evolve.

Speciation by splitting of pre-existing species

5 marks

Describe the changes that occur in gene pools during speciation.

Reproductive isolation and differential selection

7 marks

Explain how isolation leads to speciation.