B3.3 Muscle and motility [HL only]

Muscle and motility connect sarcomere contraction, antagonistic muscles, skeletons, synovial joints and locomotor adaptations to movement across biological scales efficiently.

Syllabus
First assessment 2025
Topic
B3.3
Level
HL

Sessile, motile and locomotion

HL only
A side-by-side visual showing a motile animal moving its whole body and a sessile organism still showing local movement or growth toward a stimulus.

Movement is universal in living organisms, but locomotion is movement of the whole organism from one place to another. Motile organisms locomote; sessile organisms remain attached yet can still move parts or grow directionally.

A motile animal can change location using metabolic energy and structures such as legs, wings or fins. A sessile coral can move tentacles, and a rooted plant can grow or bend toward a stimulus without locomoting.

Locomotion can improve survival and reproduction: a blackbird forages for food, a hare escapes a predator, an orangutan searches for a mate, and a whale migrates between feeding and breeding areas.

A plant shoot curving toward light demonstrates movement through differential growth, whereas a bird flying to a feeding site demonstrates locomotion because its whole body changes location.

Sessile does not mean incapable of movement, and locomotion is not cost-free: it requires energy and often increases nutritional demand and exposure to risk.

Movement adaptations

HL only

Assessment in practice

2 marks
How it is assessed

This objective is assessed through structured response.

What earns marks

Build the answer around this relationship: Euglena use a flagellum for locomotion.

Watch for

Confusing movement of cells or body parts with whole-organism locomotion.

Representative question

Question 1

[Maximum number: 2]

Microscopic eukaryotes include Euglena and Paramecium. Outline the range of cellular structures used for locomotion in these organisms.

Sliding Filaments Shorten a Sarcomere

HL only

A sarcomere extends between two Z lines. Thin actin filaments anchored at the Z lines slide past central thick myosin filaments, increasing overlap and shortening the sarcomere without shortening either filament.

Calcium released from the sarcoplasmic reticulum binds troponin, changing its shape and moving tropomyosin away from myosin-binding sites on actin. Energized myosin heads can then form cross-bridges.

Cross-bridge cycle: myosin-ADP-Pi binds actin → Pi and ADP release drives the power stroke → ATP binding detaches myosin → ATP hydrolysis re-cocks the head. Cycling continues while calcium and ATP are available.

During contraction the Z lines move closer, the I band and H zone narrow, and the A band stays the same length because thick myosin filament length is unchanged.

ATP does not directly pull actin: it permits detachment and re-cocking, while the myosin power stroke generates force. Actin and myosin slide; they do not shrink.

Sequence Contraction

HL only
Ordered contraction states.

Skeletal muscle contraction occurs when myosin heads repeatedly pull actin filaments towards the centre of each sarcomere. The filaments slide past one another; neither filament becomes shorter.

  1. Expose the binding sites: calcium ions bind to troponin, shifting tropomyosin away from binding sites on actin.
  2. Pull: a myosin head attaches to an exposed actin site. Release of ADP and phosphate during the power stroke changes the head angle and pulls actin towards the sarcomere centre.
  3. Detach: ATP binds to myosin, causing the head to detach from actin.
  4. Reset: ATP is hydrolysed to ADP and phosphate, re-cocking the detached head so it can attach again.
  5. While calcium remains available and ATP is supplied, repeated cycles increase actin and myosin overlap, bring the Z-lines closer and shorten the sarcomere.

ATP does not pull actin directly: it enables myosin detachment and re-cocking, while the power stroke of an attached myosin head produces the pull. Sliding shortens the sarcomere, not the actin or myosin filaments themselves.

Sliding filament model

HL only

Assessment in practice

1–3 marks
How it is assessed

This objective is assessed through structured response, commonly using Explain / Draw / Identify.

Command terms

Explain / Draw / Identify / Describe / Analyse / State / Outline / Label / Deduce

What earns marks

Build the answer around this relationship: Sarcomeres are repeating contractile units between Z lines.

Watch for

Saying actin or myosin filaments shorten instead of sliding past each other.

Representative question

Question 1

[Maximum number: 8]

Actin and myosin are two proteins found in muscles. Explain how skeletal muscle contracts, including the interaction of these proteins.

Trigger and reset a muscle

HL only
A compact diagram connecting motor neuron, neuromuscular junction, motor end plate, sarcoplasmic reticulum calcium release, and a titin spring inside one sarcomere.

A motor neuron triggers skeletal muscle contraction at the neuromuscular junction. An action potential reaches the motor end plate, acetylcholine is released and the sarcolemma is depolarized; the signal causes the sarcoplasmic reticulum to release Ca²⁺. Ca²⁺ exposes actin binding sites and cross-bridge cycling shortens the sarcomere. Acetylcholinesterase removes acetylcholine and Ca²⁺ pumps return calcium to the sarcoplasmic reticulum, so tropomyosin covers the binding sites and the fibre relaxes. A motor unit is one motor neuron plus all the fibres it controls. Titin provides elastic recoil, centres myosin and limits overstretching; antagonistic muscles are needed because muscle contraction produces force in one direction, not active extension.

  • One motor neuron and its muscle fibres form a motor unit.
  • Acetylcholine and Ca²⁺ initiate contraction.
  • Acetylcholinesterase and Ca²⁺ reuptake allow relaxation.
  • Titin recoils the sarcomere; antagonistic pairs reverse movement.

Titin and antagonistic muscles

HL only

Assessment in practice

1–2 marks
How it is assessed

This objective is assessed through structured response, commonly using Label / Identify / Outline.

Command terms

Label / Identify / Outline / Describe / Explain

What earns marks

Build the answer around this relationship: Titin helps keep myosin centred in the sarcomere.

Watch for

Saying antagonistic muscles both contract together for the same movement.

Representative question

Question 1

[Maximum number: 2]

Explain the role of the protein titin in muscle relaxation.

Motor units

HL only

Assessment in practice

1 marks
How it is assessed

This objective is assessed through multiple choice, commonly using Identify.

Command terms

Identify

What earns marks

Build the answer around this relationship: An action potential at the motor end plate causes neurotransmitter release.

Representative question

Question 1

[Maximum number: 1]

What happens when an action potential reaches motor end plates?

A

Calcium ions are absorbed by the muscle fibres.

B

The sarcomeres relax.

C

Neurotransmitter is released.

D

Action potential is passed to the neuron.

Skeletons Turn Muscle Pull into Useful Leverage

HL only

Skeletons provide rigid muscle anchorage and jointed levers that convert muscle tension into movement; vertebrates use endoskeletons and arthropods use exoskeletons.

In vertebrates, tendons attach muscles to bones on the outside of an internal skeleton. A joint acts as the fulcrum, muscle pull supplies the effort and a body part or external object provides the load.

An arthropod's chitinous exoskeleton surrounds the body; muscles attach to its inner surface across flexible joints. In both designs, antagonistic muscles pull on opposite sides because muscles generate force only by contracting.

At a human elbow, biceps tension transmitted by its tendon rotates the forearm around the joint. In an insect leg, internally attached flexor and extensor muscles rotate neighbouring exoskeleton plates around a joint.

Skeletons do more than support: their geometry trades force for speed or movement range. An exoskeleton is external and is not made of vertebrate bone.

Skeletons as anchorage and levers

HL only

Assessment in practice

1–2 marks
How it is assessed

This objective is assessed through structured response, commonly using State / Identify / Outline.

Command terms

State / Identify / Outline / Distinguish

What earns marks

Build the answer around this relationship: Tendons attach muscles to bones.

Watch for

Confusing tendons with ligaments when describing muscle attachment.

Representative question

Question 1

[Maximum number: 1]

Outline how the muscle attachment of insects differs from humans.

Synovial joints and range of motion

HL only
Labelled knee anatomy showing bones, cartilage, menisci, ligaments and tendons around a synovial joint.

A synovial joint permits movement while limiting friction and instability. At the hip, the head of the femur fits into the pelvis; articular cartilage covers the bone ends, a synovial membrane produces lubricating fluid, ligaments stabilize the joint, and tendons attach muscles to bones. A ball-and-socket joint such as the hip allows movement in several planes, including circumduction. A hinge joint such as the knee mainly allows flexion and extension. Range of motion can be measured as a joint angle with a goniometer or image analysis.

  • Cartilage reduces friction; synovial fluid lubricates.
  • Ligaments connect and stabilize bones; tendons connect muscle to bone.
  • Hip = ball-and-socket with wide multi-plane movement.
  • Knee = hinge joint, mainly flexion and extension.
  • Range of motion is recorded as an angle in degrees.

Synovial joint movement

HL only

Assessment in practice

1 marks
How it is assessed

This objective is assessed through structured response, commonly using State / Identify / Compare.

Command terms

State / Identify / Compare / Label / Distinguish

What earns marks

Build the answer around this relationship: Synovial fluid lubricates joints and reduces friction.

Watch for

Mixing up ligaments, which connect bone to bone, with tendons, which connect muscle to bone.

Representative question

Question 1

[Maximum number: 2]

State the function of structures I and II.

I:

II:

Range of motion

HL only

Assessment in practice

1–3 marks
How it is assessed

This objective is assessed through structured response, commonly using Compare / Identify / Describe.

Command terms

Compare / Identify / Describe

What earns marks

Build the answer around this relationship: Ball-and-socket joints allow movement in multiple planes.

Watch for

Saying the knee allows all-plane movement like the hip.

Representative question

Question 1

[Maximum number: 2]

Compare the movements of the hip joint and the knee joint.

Intercostal Muscles Change Thoracic Volume

HL only

External and internal intercostal muscles form antagonistic layers with different fibre orientations, so their contraction moves the ribcage in opposite directions.

External intercostals contract to lift ribs up and out during inspiration, increasing thoracic volume. Internal intercostals contract during forced expiration to pull ribs down and in, decreasing thoracic volume.

When one intercostal layer contracts, the opposing layer is stretched. Stretching its sarcomeres stores elastic potential energy in titin; titin recoil assists return as the active layer relaxes.

Forced expiration: internal intercostals contract → ribs move down/in → thoracic volume falls → pressure rises above atmospheric pressure → air flows out; the external layer is stretched.

Quiet expiration mainly uses relaxation and elastic recoil; strong internal-intercostal contraction is associated with forced expiration. The muscles change pressure indirectly by moving the ribs.

Marine Mammals Streamline Movement in Water

HL only

Marine mammals are adapted for efficient swimming and periodic air breathing through streamlined bodies, modified limbs, tail flukes and specialized airways.

A smooth tapered body and reduced external projections lower drag. Forelimbs form flippers for steering and control, while reduced hind limbs and pelvic structures reduce resistance in cetaceans.

Cetacean tail flukes provide propulsion by moving up and down, unlike the side-to-side tail movement typical of fish. Powerful axial muscles drive this oscillation.

A whale surfaces with its dorsal blowhole exposed, exhales and inhales rapidly, then closes the airway during the next dive; this allows periodic breathing without lifting the whole head far from the water.

Marine mammals still use lungs and must surface for air. Flippers mainly steer and stabilize in cetaceans; the tail fluke supplies the main propulsive force.

Muscle And Motility

HL only

A strong answer links movement benefit, muscle contraction mechanism, force transfer, joint range, and locomotion adaptations. For contraction, use calcium-troponin-tropomyosin and ATP-driven myosin cross-bridge cycling. For movement, use antagonistic muscles, tendons, ligaments, skeletons as levers, and joint type. For locomotion, link body form to survival or swimming advantage. Locomotion can improve survival and reproductive success.

  • Contraction answers need calcium, actin binding sites, myosin cross-bridges, ATP, and sarcomere shortening.
  • Movement answers need antagonistic pairs because muscles contract but do not actively extend.
  • Joint and skeleton answers need tendon versus ligament, lever action, synovial fluid, and range of motion.
  • Locomotion answers should link examples to food, escape, mate finding, or migration: blackbirds feeding, hares escaping predators, orangutans finding mates, and whales migrating.

Objective notes

10 learning objectives
B3.3.1(HL)—Movement adaptations• Movement is universal, but only some organisms show locomotion• Motile organisms move for food, mates, migration, or escape• Sessile organisms such as coral or plants still move body parts or grow toward stimuli1% of analysed papers 1 paper · 1 questionViewB3.3.2(HL)—Sliding filament model• Sarcomeres contain actin thin filaments and myosin thick filaments between Z lines• Calcium binds troponin, moving tropomyosin away from actin binding sites• Myosin heads use ATP in a cross-bridge cycle to slide actin and shorten sarcomeres26% of analysed papers 29 papers · 30 questionsViewB3.3.3(HL)—Titin and antagonistic muscles• Titin acts as a molecular spring in sarcomeres and centres myosin• Titin recoils after stretching and prevents overstretching• Antagonistic muscles are needed because muscles actively contract but do not actively extend8% of analysed papers 9 papers · 10 questionsViewB3.3.4(HL)—Motor units• Motor neurons connect to skeletal muscle fibres at neuromuscular junctions• Acetylcholine release at motor end plates triggers sarcolemma excitation• Excitation releases calcium from sarcoplasmic reticulum to start contraction1% of analysed papers 1 paper · 1 questionViewB3.3.5(HL)—Skeletons as anchorage and levers• Skeletons provide support, protection, muscle anchorage, and lever systems• Vertebrate endoskeletons attach muscles by tendons• Arthropod exoskeletons use chitin, joints, and internal muscle attachments4% of analysed papers 5 papers · 5 questionsViewB3.3.6(HL)—Synovial joint movement• Synovial joints contain lubricating synovial fluid between cartilage-covered bones• Tendons attach muscles to bones; ligaments connect bones and stabilize joints• The hip is a ball-and-socket joint between femur and pelvis7% of analysed papers 8 papers · 8 questionsViewB3.3.7(HL)—Range of motion• Range of motion depends on joint type, bone surfaces, ligaments, and muscles• Ball-and-socket joints such as shoulder and hip allow circumduction• Hinge joints such as elbow and knee mainly allow flexion and extension3% of analysed papers 3 papers · 3 questionsViewB3.3.8(HL)—Intercostal muscles• External and internal intercostal muscles act antagonistically during ventilation• External intercostals lift ribs up and out for inspiration• Internal intercostals pull ribs down and in during forced expiration0% of analysed papers ViewB3.3.9(HL)—Reasons for locomotion• Locomotion allows animals to forage, avoid predators, find mates, and migrate• Examples include blackbirds feeding, hares escaping predators, orangutans finding mates, and whales migrating• Locomotion can improve survival and reproductive success0% of analysed papers ViewB3.3.10(HL)—Swimming adaptations in marine mammals• Marine mammals have streamlined bodies, reduced drag, flippers, and up-down tail flukes• Reduced pelvic bones and absent hind limbs reduce resistance in water• Cetacean blowholes allow rapid surface breathing and close during dives0% of analysed papers View